Abstract

Vertebrate fossils are extremely rare below the Achanarras fish beds and equivalent strata in northern Scotland. Here we describe the cheiracanthid acanthodians from the lowest Middle Devonian of this region, comprising partial articulated specimens and squamation patches of two species Cheiracanthus flabellicostatus and C. brevicostatus. Both species were previously only known as isolated scales from the eastern Baltic and Russia. The stratigraphic range of the two species in Scotland extends up into the Achanarras equivalent fish beds in the Moray Firth.